Newer
Older
Digital_Repository / Misc / Mass downloads / UTas / 2518.html
<!DOCTYPE html PUBLIC "-//W3C//DTD XHTML 1.0 Transitional//EN"
"http://www.w3.org/TR/xhtml1/DTD/xhtml1-transitional.dtd">
<html>
  <head>
    <title>UTas ePrints - The association between substrate variability and bryophyte and lichen diversity in eastern Australian forests</title>
    <script type="text/javascript" src="http://eprints.utas.edu.au/javascript/auto.js"><!-- padder --></script>
    <style type="text/css" media="screen">@import url(http://eprints.utas.edu.au/style/auto.css);</style>
    <style type="text/css" media="print">@import url(http://eprints.utas.edu.au/style/print.css);</style>
    <link rel="icon" href="/images/eprints/favicon.ico" type="image/x-icon" />
    <link rel="shortcut icon" href="/images/eprints/favicon.ico" type="image/x-icon" />
    <link rel="Top" href="http://eprints.utas.edu.au/" />
    <link rel="Search" href="http://eprints.utas.edu.au/cgi/search" />
    <meta content="Pharo, E.J." name="eprints.creators_name" />
<meta content="Beattie, A.J." name="eprints.creators_name" />
<meta content="Emma.Pharo@utas.edu.au" name="eprints.creators_id" />
<meta content="" name="eprints.creators_id" />
<meta content="article" name="eprints.type" />
<meta content="2007-11-22 03:12:03" name="eprints.datestamp" />
<meta content="2008-01-08 15:30:00" name="eprints.lastmod" />
<meta content="show" name="eprints.metadata_visibility" />
<meta content="The association between substrate variability and bryophyte and lichen diversity in eastern Australian forests" name="eprints.title" />
<meta content="pub" name="eprints.ispublished" />
<meta content="270000" name="eprints.subjects" />
<meta content="restricted" name="eprints.full_text_status" />
<meta content="bryophyte, lichen, alpha diversity, beta diversity, fire" name="eprints.keywords" />
<meta content="The difference in species diversity on different substrates and the effect of both substrate (ground, logs, rocks, trees and fallen branches) and site variation (such as disturbance history, aspect, slope) on bryophyte and lichen diversity were investigated in eastern Australian forests. Thirty-five sites in forest types ranging from dry sclerophyll to riparian were surveyed using a 50 m x 20 m area. Despite strong differences between the different substrates in species richness and composition for both bryophytes and lichens, differences within each substrate were limited. On examining each substrate separately, we found that variation in substrate quality, such as degree of log decay, was not strongly correlated with species diversity, explaining no more than 16% of richness and 5% of composition. Despite both bryophytes and lichens showing high fidelity for particular substrates, the quality of that substrate was not an important factor in determining species diversity in this study. Site environmental variables explained larger variation in both bryophyte and lichen species diversity, in particular species richness, with individual site variables explaining up to 41% of richness (topographic position) and 6% of composition (time since fire). For some site variables, notably some of the disturbance variables, there were no overall trends, but significant results for particular substrates, such as logging being significant only on logs and trees." name="eprints.abstract" />
<meta content="2002" name="eprints.date" />
<meta content="published" name="eprints.date_type" />
<meta content="The Bryologist" name="eprints.publication" />
<meta content="105" name="eprints.volume" />
<meta content="1" name="eprints.number" />
<meta content="11-26" name="eprints.pagerange" />
<meta content="TRUE" name="eprints.refereed" />
<meta content="0007-2745" name="eprints.issn" />
<meta content="https://mywebspace.wisc.edu/jpbennet/web/abls/" name="eprints.official_url" />
<meta content="
ADAMS, D. B. &amp; P. G. RISSER. 1971. Some factors influencing the frequency of 
bark lichens in north central Oklahoma. American Journal of Botany.  58: 752-
757.
ANDERSON, D. S., R. B. DAVIES &amp; J. A. JANSSENS. 1995. Relationships of 
bryophytes and lichens to environmental gradients. Vegetatio.  120(2): 147-
159.
ASHTON, D. H. 1986. Ecology of bryophytic communities in mature Eucalyptus 
regnans F. Muell. forest at Wallaby Creek, Victoria. Australian Journal of 
Botany.  34: 107-129.
BAILEY, R. H. 1976. Ecological aspects of dispersal and establishment in lichens. In. 
Lichenology: progress and problems (eds. D. H. Brown, D. L. Hawksworth &amp; R. H. Bailey) . Academic Press, London.
BARKHAM, J. J. 1958. Phytosociology and ecology of cryptogamic epiphytes.   Van 
Gorcum Co., Assen, Netherlands.
BARKHAM, J. P. 1992. The effects of management on the ground flora of ancient 
woodland, Brigsteer Park Wood, Cumbria, England. Biological Conservation.  60: 167-187.
BATES, J. W. 1992. Influence of chemical and physical factors on Quercus and 
Fraxinus epiphytes at Loch Sunart, western Scotland: a multivariate analysis. 
Journal of Ecology.  80: 163-179.
BELLAND, R. J. &amp; D. H. VITT. 1995. Bryophyte vegetation patterns along 
environmental gradients in continental bogs. Ecoscience.  2(4): 395-407.
BILLINGS, W. D. &amp; W. B. DREW. 1938. Bark factors affecting the distribution of 
corticolous bryophytic communities. The American Midland Naturalist.  20: 
302-330.	
BINNS, D. L. 1996. Flora survey, Morisett Forestry District, Central Region, New 
South Wales. Morisset Forestry District EIS Support Document No.3, 
Research Division, State Forests of N.S.W.
BRODO, I. M. 1973. Substrate ecology. In. The Lichens (eds. V. Ahmadjian &amp; M. E. 
Hale) pp. 401-441. Academic Press, London.
CARLETON, T. J. 1990. Variation in terricolous bryophyte and macrolichen 
vegetation along primary gradients in Canadian boreal forests. Journal of 
Vegetation Science. 1: 585-594.
CLARKE, K. R. 1993. Non-parametric multivariate analyses of changes in 
community structure. Australian Journal of Ecology.  18: 117-143.
CLEAVITT, N. 2001. Disentangling moss species limitations: The role of 
physiologically based substrate specificity for six species occurring on 
substrates with varying pH and percent organic matter. The Bryologist.  104: 59-68.
FAITH, D. P., P. R. MINCHIN &amp; L. BELBIN. 1987. Compositional dissimilarity as a 
robust measure of ecological distance. Vegetatio.  69: 57-68.
FENSHAM, R. J. &amp; H. STREIMANN. 1997. Broad landscape relations of the moss 
flora from inland dry rainforest in North Queensland, Australia. The 
Bryologist.  100: 56-64.
FORESTRY COMMISSION OF NEW SOUTH WALES. 1984. Management plan for Wyong Management Area.
GERMANO, S. R. &amp; K. C. PORTO. 1997. Ecological analysis of epixylic bryophytes 
in relation to the decomposition of the substrate (Municipality of Timbauba-
Pernambuco, Brazil). Cryptogamie, Bryalogie et Lichenologie.  18: 143-150.
GUSTAFSSON, L., A. FISKESJO, T. HALLINGBACK &amp; T. INGELOG. 1992. 
Semi-natural deciduous broadleaved woods in southern Sweden- habitat 
factors of importance to some bryophyte species. Biological Conservation.  59: 175-181.
HARNER, R. F. &amp; K. T. HARPER. 1976. The role of area, heterogeneity, and 
favorability in plant species diversity of pinyon-juniper ecosystems. Ecology.  57: 1254-1263.
IWATSUKI, Z. 1990. Origin of the New Caledonian bryophtes. Tropical Bryology.  2: 139-148.
JONSSON, P. G. &amp; P. A. ESSEEN. 1990. Treefall disturbance maintains high 
bryophyte diversity in a boreal spruce forest. Journal of Ecology.  78: 924-936.
KANTVILAS, G. &amp; P. R. MINCHIN. 1989. An analysis of epiphytic lichen 
communities in Tasmanian cool temperate rainforest. Vegetatio.  84: 99-112.
P. W. JAMES &amp; S. J. JARMAN. 1985. Macrolichens in Tasmanian 
rainforest. The Lichenologist.  17(1): 76-83.
KOOIJMAN, A. M. 1992. The decrease of rich fen bryophytes in The Netherlands. 
Biological Conservation.  59: 139-143.
KOPONEN, T. 1992. Endangered bryophytes on a global scale. Biological 
Conservation.  59: 255-258.
LAMBERT, J. D. H. &amp; P. F. MAYCOCK. 1968. The ecology of terricolous lichens of 
the northern coniferous forests of central eastern Canada. Canadian Journal of 
Botany.  46: 1043-1078.
LEE, T. D. &amp; G. H. LA ROI. 1979. Gradient analysis of bryophytes in Jasper National Park, Alberta. Canadian Journal of Botany.  57: 914-925.
MAGURRAN, A. E. 1988. Ecological diversity and its measurement.   Croom Helm, 
London.
MARINO, P. C. 1988. Coexistence on divided habitats: mosses in the family 
Splachnaceae. Annales Zoologici Fennici.  25: 89-98.
MCCARTHY, P. M. 1991. Checklist of Australian Lichens. 4  National Herbarium of 
Victoria, Melbourne.
MILES, C. J. &amp; R. E. LONGTON. 1992. Deposition of moss spores in relation to 
distance from parent gametophytes. Journal of Bryology.  17: 355-368.
MINCHIN, P. R. 1991. DECODA: database for ecological community data. 
Australian National University, Canberra.
NAGANO, I. 1969. Comparative studies of moss vegetation developing on limestone, 
chert, and other rocks lying adjacent to each other in the Chichibu Mountain 
area, central Japan. Journal of the Hattori Botanical Laboratory.  32: 155-203.
PALMER, M. W. 1986. Pattern in corticolous bryophyte communities of the North 
Carolina Piedmont: do mosses see the forest or the trees? The Bryologist.  
89(1): 59-65.
PHARO, E. J. &amp; A. J. BEATTIE. 1997. Bryophyte and lichen diversity: a comparative study. Australian Journal of Ecology.  22: 151-162.
PHARO, E. J. &amp; A. J. BEATTIE. 2001. Management forest types as a surrogate for 
vascular plant, bryophyte and lichen diversity. Australian Journal of Botany.  49: 1-8.
PHARO, E. J. &amp; D. H. VITT. 2000. Local variation in bryophyte and macro-lichen cover and diversity in montane forests of western Canada. The Bryologist.  103: 455-
466.
RAMBO, T. R. &amp; P. S. MUIR. 1998. Forest floor bryophytes of Pseudotsuga 
menziesii-Tsuga heterophylla stands in Oregon: influences of substrate and 
overstory. The Bryologist.  101: 116-130.
SCHIEFERSTEIN, B. &amp; K. LORIS. 1992. Ecological investigations on lichen fields 
of the Central Namib. Vegetatio.  98: 113-128.
SCHMITT, C. K. &amp; N. G. SLACK. 1990. Host specificity of epiphytic lichens and 
bryophytes: a comparison of the Adirondack Mountains (New York) and the 
Southern Blue Ridge Mountains (North Carolina). The Bryologist.  93(3): 257-274.
SCHULTER, D. &amp; R. E. RICKLEFS. 1993. Species diversity: an introduction to the 
problem. In. Species diversity in ecological communities: historical and 
geographical perspectives (eds. R. E. Ricklefs &amp; D. Schluter) pp. 1-10. The University of Chicago Press, Chicago.
SCOTT, G. A. M. 1970. Vegetation studies on Secretary Island, Fiordland. Part II: 
epiphytic and ground cryptogamic vegetation on the northern slopes. New 
Zealand Journal of Botany.  8: 30-50.
SCOTT, G. A. M. &amp; J. A. BRADSHAW. 1986. Australian Liverworts (Hepaticae). Brunonia.  8(1): 1-171.
SLACK, N. G. 1977. Species diversity and community structure in bryophytes: New 
York State studies. Bulletin of the New York State Museum.  428: 1-70.
SLACK, N. G. 1984. A new look at bryophyte community analysis: field and statistical methods. Journal of the Hattori Botanical Laboratory.  55: 113-132.
SÖDERSTRÖM, L. 1981. Distribution of bryophytes in spruce forests on hill slopes 
in central Sweden. Wahlenbergia.  7: 141-153.
SÖDERSTRÖM, L. 1989. Regional distribution patterns of bryophyte species on spruce logs in northern Sweden. The Bryologist.  92(3): 349-355.
SÖDERSTRÖM, L.    1993. Substrate preference in some forest bryophytes: a quantitative study. Lindbergia. A Journal of Bryology.  18(3): 98-103.
SPSS FOR WINDOWS. 1983. Release 6.0. Chicago, Illanois.
STATE FORESTS OF NEW SOUTH WALES. 1995. Morisset Forestry District 
Environmental Impact Statement. State Forests of New South Wales, Pennant 
Hills, Sydney.
STREIMANN, H. &amp; J. CURNOW. 1989. Catalogue of mosses of Australia and its 
external territories. Australian Flora and Fauna Series 10 Australian Government Publishing Service Press, Canberra.
SUREN, A. M. 1996. Bryophyte distribution patterns in relation to macro-, meso- and 
micro-scale variables in south island New Zealand streams. New Zealand 
Journal of Marine and Freshwater Research.  30: 501-523.
TANGNEY, R. S., J. B. WILSON &amp; A. F. MARK. 1990. Bryophyte island 
biogeography: a study in Lake Manapouri, New Zealand. Oikos.  59: 21-26.
TER BRAAK, C. J. F. 1991. CANOCO- a FORTRAN program for Canonical 
Community Ordination. Microcomputer Power, Ithaca, New York.
VAN ZANTEN, B. O. &amp; T. PÓCS. 1981. Distribution and dispersal of bryophytes. 
Advances in Bryology.  1: 479-562.
VITT, D. H., M. OSTAFICHUK &amp; I. BRODO. 1973. Foliicolous lichens of Thuja 
plicata in western British Colombia. Canadian Journal of Botany.  51: 571-580.
VITT, D. H., Y. LI &amp; R. J. BELLAND. 1995. Patterns of bryophyte diversity in 
peatlands of continental Western Canada. The Bryologist.  98(2): 218-227.
WOLF, J. H. D. 1994. Factors controlling the distribution of vascular and non-
vascular epiphytes in the northern Andes. Vegetatio.  112: 15-28.
ZAR, J.H., 1996. Biostatistical analysis. 3rd edition. Prentice Hall, New Jersey. 662pp." name="eprints.referencetext" />
<meta content="Pharo, E.J. and Beattie, A.J. (2002) The association between substrate variability and bryophyte and lichen diversity in eastern Australian forests. The Bryologist, 105 (1). pp. 11-26. ISSN 0007-2745" name="eprints.citation" />
<meta content="http://eprints.utas.edu.au/2518/1/Pharo_and_Beattie_2002.pdf" name="eprints.document_url" />
<link rel="schema.DC" href="http://purl.org/DC/elements/1.0/" />
<meta content="The association between substrate variability and bryophyte and lichen diversity in eastern Australian forests" name="DC.title" />
<meta content="Pharo, E.J." name="DC.creator" />
<meta content="Beattie, A.J." name="DC.creator" />
<meta content="270000 Biological Sciences" name="DC.subject" />
<meta content="The difference in species diversity on different substrates and the effect of both substrate (ground, logs, rocks, trees and fallen branches) and site variation (such as disturbance history, aspect, slope) on bryophyte and lichen diversity were investigated in eastern Australian forests. Thirty-five sites in forest types ranging from dry sclerophyll to riparian were surveyed using a 50 m x 20 m area. Despite strong differences between the different substrates in species richness and composition for both bryophytes and lichens, differences within each substrate were limited. On examining each substrate separately, we found that variation in substrate quality, such as degree of log decay, was not strongly correlated with species diversity, explaining no more than 16% of richness and 5% of composition. Despite both bryophytes and lichens showing high fidelity for particular substrates, the quality of that substrate was not an important factor in determining species diversity in this study. Site environmental variables explained larger variation in both bryophyte and lichen species diversity, in particular species richness, with individual site variables explaining up to 41% of richness (topographic position) and 6% of composition (time since fire). For some site variables, notably some of the disturbance variables, there were no overall trends, but significant results for particular substrates, such as logging being significant only on logs and trees." name="DC.description" />
<meta content="2002" name="DC.date" />
<meta content="Article" name="DC.type" />
<meta content="PeerReviewed" name="DC.type" />
<meta content="application/pdf" name="DC.format" />
<meta content="http://eprints.utas.edu.au/2518/1/Pharo_and_Beattie_2002.pdf" name="DC.identifier" />
<meta content="https://mywebspace.wisc.edu/jpbennet/web/abls/" name="DC.relation" />
<meta content="Pharo, E.J. and Beattie, A.J. (2002) The association between substrate variability and bryophyte and lichen diversity in eastern Australian forests. The Bryologist, 105 (1). pp. 11-26. ISSN 0007-2745" name="DC.identifier" />
<meta content="http://eprints.utas.edu.au/2518/" name="DC.relation" />
<link rel="alternate" href="http://eprints.utas.edu.au/cgi/export/2518/BibTeX/epprod-eprint-2518.bib" title="BibTeX" type="text/plain" />
<link rel="alternate" href="http://eprints.utas.edu.au/cgi/export/2518/ContextObject/epprod-eprint-2518.xml" title="OpenURL ContextObject" type="text/xml" />
<link rel="alternate" href="http://eprints.utas.edu.au/cgi/export/2518/ContextObject::Dissertation/epprod-eprint-2518.xml" title="OpenURL Dissertation" type="text/xml" />
<link rel="alternate" href="http://eprints.utas.edu.au/cgi/export/2518/ContextObject::Journal/epprod-eprint-2518.xml" title="OpenURL Journal" type="text/xml" />
<link rel="alternate" href="http://eprints.utas.edu.au/cgi/export/2518/DC/epprod-eprint-2518.txt" title="Dublin Core" type="text/plain" />
<link rel="alternate" href="http://eprints.utas.edu.au/cgi/export/2518/DIDL/epprod-eprint-2518.xml" title="DIDL" type="text/xml" />
<link rel="alternate" href="http://eprints.utas.edu.au/cgi/export/2518/EndNote/epprod-eprint-2518.enw" title="EndNote" type="text/plain" />
<link rel="alternate" href="http://eprints.utas.edu.au/cgi/export/2518/HTML/epprod-eprint-2518.html" title="HTML Citation" type="text/html; charset=utf-8" />
<link rel="alternate" href="http://eprints.utas.edu.au/cgi/export/2518/METS/epprod-eprint-2518.xml" title="METS" type="text/xml" />
<link rel="alternate" href="http://eprints.utas.edu.au/cgi/export/2518/MODS/epprod-eprint-2518.xml" title="MODS" type="text/xml" />
<link rel="alternate" href="http://eprints.utas.edu.au/cgi/export/2518/RIS/epprod-eprint-2518.ris" title="Reference Manager" type="text/plain" />
<link rel="alternate" href="http://eprints.utas.edu.au/cgi/export/2518/Refer/epprod-eprint-2518.refer" title="Refer" type="text/plain" />
<link rel="alternate" href="http://eprints.utas.edu.au/cgi/export/2518/Simple/epprod-eprint-2518text" title="Simple Metadata" type="text/plain" />
<link rel="alternate" href="http://eprints.utas.edu.au/cgi/export/2518/Text/epprod-eprint-2518.txt" title="ASCII Citation" type="text/plain; charset=utf-8" />
<link rel="alternate" href="http://eprints.utas.edu.au/cgi/export/2518/XML/epprod-eprint-2518.xml" title="EP3 XML" type="text/xml" />

  </head>
  <body bgcolor="#ffffff" text="#000000" onLoad="loadRoutine(); MM_preloadImages('images/eprints/ePrints_banner_r5_c5_f2.gif','images/eprints/ePrints_banner_r5_c7_f2.gif','images/eprints/ePrints_banner_r5_c8_f2.gif','images/eprints/ePrints_banner_r5_c9_f2.gif','images/eprints/ePrints_banner_r5_c10_f2.gif','images/eprints/ePrints_banner_r5_c11_f2.gif','images/eprints/ePrints_banner_r6_c4_f2.gif')">
    
    <div class="ep_noprint"><noscript><style type="text/css">@import url(http://eprints.utas.edu.au/style/nojs.css);</style></noscript></div>




<table width="795" border="0" cellspacing="0" cellpadding="0">
  <tr>
    <td><script language="JavaScript1.2">mmLoadMenus();</script>
      <table border="0" cellpadding="0" cellspacing="0" width="795">
        <!-- fwtable fwsrc="eprints_banner_final2.png" fwbase="ePrints_banner.gif" fwstyle="Dreamweaver" fwdocid = "1249563342" fwnested="0" -->
        <tr>
          <td><img src="/images/eprints/spacer.gif" width="32" height="1" border="0" alt="" /></td>
          <td><img src="/images/eprints/spacer.gif" width="104" height="1" border="0" alt="" /></td>
          <td><img src="/images/eprints/spacer.gif" width="44" height="1" border="0" alt="" /></td>
          <td><img src="/images/eprints/spacer.gif" width="105" height="1" border="0" alt="" /></td>
          <td><img src="/images/eprints/spacer.gif" width="41" height="1" border="0" alt="" /></td>
          <td><img src="/images/eprints/spacer.gif" width="16" height="1" border="0" alt="" /></td>
          <td><img src="/images/eprints/spacer.gif" width="68" height="1" border="0" alt="" /></td>
          <td><img src="/images/eprints/spacer.gif" width="68" height="1" border="0" alt="" /></td>
          <td><img src="/images/eprints/spacer.gif" width="68" height="1" border="0" alt="" /></td>
          <td><img src="/images/eprints/spacer.gif" width="82" height="1" border="0" alt="" /></td>
          <td><img src="/images/eprints/spacer.gif" width="69" height="1" border="0" alt="" /></td>
          <td><img src="/images/eprints/spacer.gif" width="98" height="1" border="0" alt="" /></td>
          <td><img src="/images/eprints/spacer.gif" width="1" height="1" border="0" alt="" /></td>
        </tr>
        <tr>
          <td colspan="12"><img name="ePrints_banner_r1_c1" src="/images/eprints/ePrints_banner_r1_c1.gif" width="795" height="10" border="0" alt="" /></td>
          <td><img src="/images/eprints/spacer.gif" width="1" height="10" border="0" alt="" /></td>
        </tr>
        <tr>
          <td rowspan="6"><img name="ePrints_banner_r2_c1" src="/images/eprints/ePrints_banner_r2_c1.gif" width="32" height="118" border="0" alt="" /></td>
          <td rowspan="5"><a href="http://www.utas.edu.au/"><img name="ePrints_banner_r2_c2" src="/images/eprints/ePrints_banner_r2_c2.gif" width="104" height="103" border="0" alt="" /></a></td>
          <td colspan="10"><img name="ePrints_banner_r2_c3" src="/images/eprints/ePrints_banner_r2_c3.gif" width="659" height="41" border="0" alt="" /></td>
          <td><img src="/images/eprints/spacer.gif" width="1" height="41" border="0" alt="" /></td>
        </tr>
        <tr>
          <td colspan="3"><a href="http://eprints.utas.edu.au/"><img name="ePrints_banner_r3_c3" src="/images/eprints/ePrints_banner_r3_c3.gif" width="190" height="31" border="0" alt="" /></a></td>
          <td rowspan="2" colspan="7"><img name="ePrints_banner_r3_c6" src="/images/eprints/ePrints_banner_r3_c6.gif" width="469" height="37" border="0" alt="" /></td>
          <td><img src="/images/eprints/spacer.gif" width="1" height="31" border="0" alt="" /></td>
        </tr>
        <tr>
          <td colspan="3"><img name="ePrints_banner_r4_c3" src="/images/eprints/ePrints_banner_r4_c3.gif" width="190" height="6" border="0" alt="" /></td>
          <td><img src="/images/eprints/spacer.gif" width="1" height="6" border="0" alt="" /></td>
        </tr>
        <tr>
          <td colspan="2"><img name="ePrints_banner_r5_c3" src="/images/eprints/ePrints_banner_r5_c3.gif" width="149" height="1" border="0" alt="" /></td>
          <td rowspan="2" colspan="2"><a href="/information.html" onMouseOut="MM_swapImgRestore();MM_startTimeout()" onMouseOver="MM_showMenu(window.mm_menu_0821132634_0,0,25,null,'ePrints_banner_r5_c5');MM_swapImage('ePrints_banner_r5_c5','','/images/eprints/ePrints_banner_r5_c5_f2.gif',1);"><img name="ePrints_banner_r5_c5" src="/images/eprints/ePrints_banner_r5_c5.gif" width="57" height="25" border="0" alt="About" /></a></td>
          <td rowspan="2"><a href="/view/" onMouseOut="MM_swapImgRestore();MM_startTimeout()" onMouseOver="MM_showMenu(window.mm_menu_0821133021_1,0,25,null,'ePrints_banner_r5_c7');MM_swapImage('ePrints_banner_r5_c7','','/images/eprints/ePrints_banner_r5_c7_f2.gif',1);"><img name="ePrints_banner_r5_c7" src="/images/eprints/ePrints_banner_r5_c7.gif" width="68" height="25" border="0" alt="Browse" /></a></td>
          <td rowspan="2"><a href="/perl/search/simple" onMouseOut="MM_swapImgRestore();MM_startTimeout()" onMouseOver="MM_showMenu(window.mm_menu_0821133201_2,0,25,null,'ePrints_banner_r5_c8');MM_swapImage('ePrints_banner_r5_c8','','/images/eprints/ePrints_banner_r5_c8_f2.gif',1);"><img name="ePrints_banner_r5_c8" src="/images/eprints/ePrints_banner_r5_c8.gif" width="68" height="25" border="0" alt="Search" /></a></td>
          <td rowspan="2"><a href="/perl/register" onMouseOut="MM_swapImgRestore();MM_startTimeout();" onMouseOver="MM_showMenu(window.mm_menu_1018171924_3,0,25,null,'ePrints_banner_r5_c9');MM_swapImage('ePrints_banner_r5_c9','','/images/eprints/ePrints_banner_r5_c9_f2.gif',1);"><img name="ePrints_banner_r5_c9" src="/images/eprints/ePrints_banner_r5_c9.gif" width="68" height="25" border="0" alt="register" /></a></td>
          <td rowspan="2"><a href="/perl/users/home" onMouseOut="MM_swapImgRestore();MM_startTimeout()" onMouseOver="MM_showMenu(window.mm_menu_0821133422_4,0,25,null,'ePrints_banner_r5_c10');MM_swapImage('ePrints_banner_r5_c10','','/images/eprints/ePrints_banner_r5_c10_f2.gif',1);"><img name="ePrints_banner_r5_c10" src="/images/eprints/ePrints_banner_r5_c10.gif" width="82" height="25" border="0" alt="user area" /></a></td>
          <td rowspan="2"><a href="/help/" onMouseOut="MM_swapImgRestore();MM_startTimeout()" onMouseOver="MM_showMenu(window.mm_menu_0821133514_5,0,25,null,'ePrints_banner_r5_c11');MM_swapImage('ePrints_banner_r5_c11','','/images/eprints/ePrints_banner_r5_c11_f2.gif',1);"><img name="ePrints_banner_r5_c11" src="/images/eprints/ePrints_banner_r5_c11.gif" width="69" height="25" border="0" alt="Help" /></a></td>
          <td rowspan="3" colspan="4"><img name="ePrints_banner_r5_c12" src="/images/eprints/ePrints_banner_r5_c12.gif" width="98" height="40" border="0" alt="" /></td>
          <td><img src="/images/eprints/spacer.gif" width="1" height="1" border="0" alt="" /></td>
        </tr>
        <tr>
          <td rowspan="2"><img name="ePrints_banner_r6_c3" src="/images/eprints/ePrints_banner_r6_c3.gif" width="44" height="39" border="0" alt="ePrints home" /></td>
          <td><a href="/" onMouseOut="MM_swapImgRestore()" onMouseOver="MM_swapImage('ePrints_banner_r6_c4','','/images/eprints/ePrints_banner_r6_c4_f2.gif',1);"><img name="ePrints_banner_r6_c4" src="/images/eprints/ePrints_banner_r6_c4.gif" width="105" height="24" border="0" alt="ePrints home" /></a></td>
          <td><img src="/images/eprints/spacer.gif" width="1" height="24" border="0" alt="" /></td>
        </tr>
        <tr>
          <td><img name="ePrints_banner_r7_c2" src="/images/eprints/ePrints_banner_r7_c2.gif" width="104" height="15" border="0" alt="" /></td>
          <td colspan="8"><img name="ePrints_banner_r7_c4" src="/images/eprints/ePrints_banner_r7_c4.gif" width="517" height="15" border="0" alt="" /></td>
          <td><img src="/images/eprints/spacer.gif" width="1" height="15" border="0" alt="" /></td>
        </tr>
      </table></td>
  </tr>
    <tr><td><table width="100%" style="font-size: 90%; border: solid 1px #ccc; padding: 3px"><tr>
      <td align="left"><a href="http://eprints.utas.edu.au/cgi/users/home">Login</a> | <a href="http://eprints.utas.edu.au/cgi/register">Create Account</a></td>
      <td align="right" style="white-space: nowrap">
        <form method="get" accept-charset="utf-8" action="http://eprints.utas.edu.au/cgi/search" style="display:inline">
          <input class="ep_tm_searchbarbox" size="20" type="text" name="q" />
          <input class="ep_tm_searchbarbutton" value="Search" type="submit" name="_action_search" />
          <input type="hidden" name="_order" value="bytitle" />
          <input type="hidden" name="basic_srchtype" value="ALL" />
          <input type="hidden" name="_satisfyall" value="ALL" />
        </form>
      </td>
    </tr></table></td></tr>
  <tr>
    <td class="toplinks"><!-- InstanceBeginEditable name="content" -->


<div align="center">
  
  <table width="720" class="ep_tm_main"><tr><td align="left">
    <h1 class="ep_tm_pagetitle">The association between substrate variability and bryophyte and lichen diversity in eastern Australian forests</h1>
    <p style="margin-bottom: 1em" class="not_ep_block"><span class="person_name">Pharo, E.J.</span> and <span class="person_name">Beattie, A.J.</span> (2002) <xhtml:em>The association between substrate variability and bryophyte and lichen diversity in eastern Australian forests.</xhtml:em> The Bryologist, 105 (1). pp. 11-26. ISSN 0007-2745</p><p style="margin-bottom: 1em" class="not_ep_block"></p><table style="margin-bottom: 1em" class="not_ep_block"><tr><td valign="top" style="text-align:center"><a href="http://eprints.utas.edu.au/2518/1/Pharo_and_Beattie_2002.pdf"><img alt="[img]" src="http://eprints.utas.edu.au/style/images/fileicons/application_pdf.png" class="ep_doc_icon" border="0" /></a></td><td valign="top"><a href="http://eprints.utas.edu.au/2518/1/Pharo_and_Beattie_2002.pdf"><span class="ep_document_citation">PDF</span></a> - Full text restricted - Requires a PDF viewer<br />1886Kb</td><td><form method="get" accept-charset="utf-8" action="http://eprints.utas.edu.au/cgi/request_doc"><input accept-charset="utf-8" value="3315" name="docid" type="hidden" /><div class=""><input value="Request a copy" name="_action_null" class="ep_form_action_button" onclick="return EPJS_button_pushed( '_action_null' )" type="submit" /> </div></form></td></tr></table><p style="margin-bottom: 1em" class="not_ep_block">Official URL: <a href="https://mywebspace.wisc.edu/jpbennet/web/abls/">https://mywebspace.wisc.edu/jpbennet/web/abls/</a></p><div class="not_ep_block"><h2>Abstract</h2><p style="padding-bottom: 16px; text-align: left; margin: 1em auto 0em auto">The difference in species diversity on different substrates and the effect of both substrate (ground, logs, rocks, trees and fallen branches) and site variation (such as disturbance history, aspect, slope) on bryophyte and lichen diversity were investigated in eastern Australian forests. Thirty-five sites in forest types ranging from dry sclerophyll to riparian were surveyed using a 50 m x 20 m area. Despite strong differences between the different substrates in species richness and composition for both bryophytes and lichens, differences within each substrate were limited. On examining each substrate separately, we found that variation in substrate quality, such as degree of log decay, was not strongly correlated with species diversity, explaining no more than 16% of richness and 5% of composition. Despite both bryophytes and lichens showing high fidelity for particular substrates, the quality of that substrate was not an important factor in determining species diversity in this study. Site environmental variables explained larger variation in both bryophyte and lichen species diversity, in particular species richness, with individual site variables explaining up to 41% of richness (topographic position) and 6% of composition (time since fire). For some site variables, notably some of the disturbance variables, there were no overall trends, but significant results for particular substrates, such as logging being significant only on logs and trees.</p></div><table style="margin-bottom: 1em" cellpadding="3" class="not_ep_block" border="0"><tr><th valign="top" class="ep_row">Item Type:</th><td valign="top" class="ep_row">Article</td></tr><tr><th valign="top" class="ep_row">Keywords:</th><td valign="top" class="ep_row">bryophyte, lichen, alpha diversity, beta diversity, fire</td></tr><tr><th valign="top" class="ep_row">Subjects:</th><td valign="top" class="ep_row"><a href="http://eprints.utas.edu.au/view/subjects/270000.html">270000 Biological Sciences</a></td></tr><tr><th valign="top" class="ep_row">ID Code:</th><td valign="top" class="ep_row">2518</td></tr><tr><th valign="top" class="ep_row">Deposited By:</th><td valign="top" class="ep_row"><span class="ep_name_citation"><span class="person_name">Dr Emma Pharo</span></span></td></tr><tr><th valign="top" class="ep_row">Deposited On:</th><td valign="top" class="ep_row">22 Nov 2007 14:12</td></tr><tr><th valign="top" class="ep_row">Last Modified:</th><td valign="top" class="ep_row">09 Jan 2008 02:30</td></tr><tr><th valign="top" class="ep_row">ePrint Statistics:</th><td valign="top" class="ep_row"><a target="ePrintStats" href="/es/index.php?action=show_detail_eprint;id=2518;">View statistics for this ePrint</a></td></tr></table><p align="right">Repository Staff Only: <a href="http://eprints.utas.edu.au/cgi/users/home?screen=EPrint::View&amp;eprintid=2518">item control page</a></p>
  </td></tr></table>
</div>



    <!-- InstanceEndEditable --></td>
  </tr>
  <tr>
    <td><!-- #BeginLibraryItem "/Library/footer_eprints.lbi" -->
    <table width="795" border="0" align="left" cellpadding="0" class="footer">
  <tr valign="top">
<td colspan="2"><div align="center"><a href="http://www.utas.edu.au">UTAS home</a> | <a href="http://www.utas.edu.au/library/">Library home</a> | <a href="/">ePrints home</a> | <a href="/contact.html">contact</a> | <a href="/information.html">about</a> | <a href="/view/">browse</a> | <a href="/perl/search/simple">search</a> | <a href="/perl/register">register</a> | <a href="/perl/users/home">user area</a> | <a href="/help/">help</a></div><br /></td>
</tr>
<tr><td colspan="2"><p><img src="/images/eprints/footerline.gif" width="100%" height="4" /></p></td></tr>
      <tr valign="top">
        <td width="68%" class="footer">Authorised by the University Librarian<br />
© University of Tasmania ABN 30 764 374 782<br />
      <a href="http://www.utas.edu.au/cricos/">CRICOS Provider Code 00586B</a> | <a href="http://www.utas.edu.au/copyright/copyright_disclaimers.html">Copyright &amp; Disclaimers</a> | <a href="http://www.utas.edu.au/accessibility/index.html">Accessibility</a> | <a href="http://eprints.utas.edu.au/feedback/">Site Feedback</a>  </td>
        <td width="32%"><div align="right">
            <p align="right" class="NoPrint"><a href="http://www.utas.edu.au/"><img src="http://www.utas.edu.au/shared/logos/unioftasstrip.gif" alt="University of Tasmania Home Page" width="260" height="16" border="0" align="right" /></a></p>
            <p align="right" class="NoPrint"><a href="http://www.utas.edu.au/"><br />
            </a></p>
        </div></td>
      </tr>
      <tr valign="top">
        <td><p>  </p></td>
        <td><div align="right"><span class="NoPrint"><a href="http://www.eprints.org/software/"><img src="/images/eprintslogo.gif" alt="ePrints logo" width="77" height="29" border="0" align="bottom" /></a></span></div></td>
      </tr>
    </table>
    <!-- #EndLibraryItem -->
    <div align="center"></div></td>
  </tr>
</table>

  </body>
</html>